Postdoctoral Research Associate
Ecology and Evolutionary Biology
Princeton University, New Jersey, USA
2021-2023
The Nneji Ecology, Genetics, Evolution and Conservation (NEGEC) Lab studies the diversity and distribution of amphibians, reptiles and fish in West and Central Africa using combined field surveys, morphological, ecological and genetic data to improve understanding of their ecology, adaptation patterns and conservation status. The NEGEC lab seeks to understand the evolutionary processes that govern the diversification of West and Central African wildlife and the impact of human activities and climate change on their diversity and distribution. Also, we are interested in understanding the feeding ecology of West and Central African mammals. We welcome students, postdocs, and collaborators interested in evolution, systematics, biogeography, phylogenetics, comparative genomics and trophic ecology of West and Central African fauna. We are dedicated to communicating science to a broad audience of students, teachers, and the public through conservation outreach programs in our study areas in West and Central Africa. Additionally, the NEGEC lab engages in citizen science, where we engage local community members in scientific data collection for research and conservation planning.
Ecology and Evolutionary Biology
Princeton University, New Jersey, USA
2021-2023
Herpetology
Kunming Institute of Zoology, Chinese Academy of Sciences, China
2018-2020
Genetics
University of Chinese Academy of Sciences, China
2015-2018
Zoology (Ecology and Environmental Biology)
University of Ibadan, Nigeria
2013-2015
Biology
University of Abuja, Nigeria
2006-2011
Science Laboratory Technology
Federal Polytechnic Oko, Nigeria
2002-2004
2025: PADI Foundation Grant: Comparison of species-specific qPCR and metabarcoding methods to detect distribution patterns of highly threatened freshwater stingrays in Nigeria from environmental DNA.
2025: Mohammed bin Zayed Species Conservation: Saving the last remaining populations of the critically endangered Niger stingray (genus Fontitrygon) in Nigeria through research and actions.
2024: Conservation Nation Firefly Grant: Conservation education across local schools in Obudu Plateau, Nigeria.
2024: Rufford Foundation: Saving the last remaining populations of the highly threatened Nigerian montane herpetofauna through conservation education and community awareness programmes.
2023-2025: National Geographic Grant: Scaling Up Conservation of Evolutionarily Distinct and Globally Endangered (EDGE) Amphibians on Obudu Plateau, Nigeria
2023: Meridian National Geographic Grant: ‘Developing Multidisciplinary Approach towards sustainable conservation of the threatened Nigerian-Cameroon highlands’
2022: First Booster Rufford Small Grant: ‘Enhancing studies and sustainable conservation actions for amphibians and reptiles in highly biodiverse and threatened Nigerian montane regions’
2022: Emerging Conservationist Grant by the Conservation Nation for the project ‘Saving the Critically Endangered and Endemic Toad in Nigeria’.
2022: Future Conservationist Award by the Conservation Leadership Programme for the project ‘Conservation status of a poorly known and endemic lizard (Ondo Forest gecko) from Nigeria’
2021-24 : International Scientific Coordination Network - South (GDRI-Sud) to support ‘West African Marine Fish DNA Barcoding Network (WAMBA-net)’.
Nneji, L. M., Oladipo, S. O., Nneji, I. C., Atofarati, O. T., Asiamah, M. K., & Adelakun, K. M. (2025). Evaluating eDNA metabarcoding for fish biodiversity assessment in Nigerian aquatic ecosystems: potential, limitations, and comparisons with traditional methods. Journal of Freshwater Ecology, 40(1). https://doi.org/10.1080/02705060.2025.2541689.
Oladipo, S., Everett, A., Durand, J.-D., Adelakun, K. M., Dieudonne, W. P., Maame, A., Nneji, I., Ayoola, A. O., Atofarati, O. T., Kachi, J. B., Nneji, L. (2025). Morphology, DNA Barcoding and Range Extension of a Poorly Known Freshwater Stingray Fontitrygon garouaensis Stauch & Blanc, 1962 from Nigerian Inland Water. Integrative and Comparative Biology. https://doi.org/10.1093/icb/icaf125.
Oladipo, S., Smith, K., Everett, A., Adelakun, K. M., Nneji, L. (2025). An updated ichthyofaunal checklist for the Nigerian Lagos Lagoon with notes on nomenclatural changes and the conservation status of species. Zootaxa, 5646(1), 38–62.
Efenakpo, O. D., Nneji, L., Luiselli, L., Adeyi, A. O., Eniang, E. A., Akani, G. C., Battisti, C., Agbons, A. M. (2025). Human–reptile cohabitation and the associated conservation dilemmas in Nigeria. Biodiversity.
Zuo, B., Chen, R.-M., Tang, X.-L., Shao, Y., Liu, X.-L., Nneji, L., Sun, Y.-B. (2024). Genomic Insights Into Genetic Basis of Evolutionary Conservatism and Innovation in Frogs. Integrative Zoology. https://onlinelibrary.wiley.com/doi/10.1111/1749-4877.12931.
Kosch, T.A., Torres-Sánchez, M., Liedtke, H.C. … the Amphibian Genomics Consortium (Nneji, L. M.* is a contributing member of the Amphibian Genomics Consortium) (2024). The Amphibian Genomics Consortium: advancing genomic and genetic resources for amphibian research and conservation. BMC Genomics 25, 1025 (2024). https://doi.org/10.1186/s12864-024-10899-7.
Wu, Y.-H., Xu, M.-H., Suwannapoom, C., Nguyen, S. N., Murphy, R. W., Papenfuss, T. J., Lathrop, A., Kilunda, F. K., Gao, W., Yuan, Z.-Y., Chen, J.-M., Zhang, L., Zhao, H.-P., Wang, L.-J., Rahman, M. M., Nneji, L. M., Zhao, G.-G., Wang, Y.-Y., Jin, J.-Q., Zhang, P., Che, J. (2024). A comprehensive molecular phylogeny of the genus Sylvirana (Anura: Ranidae) highlights unrecognized diversity, revised classification and historical biogeography. Molecular Phylogenetics and Evolution. https://doi.org/10.1016/j.ympev.2024.108223.
Oladipo, S., Ayoola, A. O., Ross, K., Edah, B., Adelakun, K. M., Mustapha, M., Carine Adous, C., Gourène, B., Diop, M., Kamla, A. T., Dieudonne, W. P., Nneji, I. C., Akinsemolu, A. A., Durand, J.-D., Nneji, L. (2024). DNA Barcoding and Morphological Characterization of Marine Stingrays (Genus Fontitrygon) from the African Gulf of Guinea. Aquatic Conservation: Marine and Freshwater Ecosystems, 34(7), e4231. https://doi.org/10.1002/aqc.4231.
Oyebanji, O. O., Onditi, K. O., Azevedo, J. A., Rahaingoson, F. R., Nneji, L., Adeleye, M. A., Stull, G. W., Zhang, R., Yi, T.-S. (2023). Biogeographic patterns and environmental drivers of species richness in the globally distributed Millettioid/Phaseoloid clade (Fabaceae, subfamily Papilionoideae). Frontiers in Ecology and Evolution, 11. http://dx.doi.org/10.3389/fevo.2023.1231553
Nneji, L., Azevedo, J. A., Oyebanji, O. O., Ma, L., Elsen, P. R., Oladipo, S. O., Salako, G., Puschendorf, R., Pringle, R. M. (2023). Patterns of species richness and turnover in endemic amphibians of the Guineo‐Congolian rain forest. Diversity and Distributions, 29(8), 1035-1051. http://dx.doi.org/10.1111/ddi.13717.
Zuo, B., Nneji, L., Sun, Y.-B. (2023). Comparative genomics reveals insights into anuran genome size evolution. BMC Genomics, 24(1). http://dx.doi.org/10.1186/s12864-023-09499-8.
Adeola, A. C., Bello, S., Abdussamad, A., Mark, A., Sanke, O., Onoja, A., Nneji, L., Abdullahi, N., Olaogun, S., Rogo, L., Mangbon, G., Pedro, S., Hiinan, M., Mukhtar, M., Ibrahim, J., Saidu, H., Dawuda, P., Bala, R., Abdullahi, H., Salako, A., Kdidi, S., Yahyaoui, M. H., Yin, T.-T. (2023). Polymorphism of prion protein gene (PRNP) in Nigerian sheep. Prion, 17(1), 44-54.
Babarinde, I., Adeola, A. C., Djagoun, C. A. M. S., Nneji, L., Okeyoyin, A. O., Niba, G., Wanzie, N., Oladipo, O. C., Adebambo, A., Bello, S., Ng’ang’a, S., Olaniyi, W., Okoro, V., Adedeji, B., Olatunde, O., Ayoola, A. O., Matouke, M., Wang, Y.-y., Sanke, O., Oseni, S., Nwani, C., Murphy, R. (2023). Population structure and evolutionary history of the greater cane rat (Thryonomys swinderianus) from the Guinean Forests of West Africa. Frontiers in Genetics, 14. https://doi.org/10.3389/fgene.2023.1041103.
Xie, H.-B., Yan, C., Adeola, A., Wang, K., Haung, C.-P., Xu, M.-M., Qiu, Q., Yin, X., Fan, C.-Y., Ma, Y.-F., Yin, T.-T., Gao, Y., Deng, J.-K., Okeyoyin, A., Oluwole, O., Omotoso, O., Okoro, V., Omitogun, O., Dawuda, P., Olaogun, S., Nneji, L., Ayoola, A., Sanke, O., Luka, P., Okoth, E., Lekelool, I., Mijele, D., Bishop, R., Han, J., Wang, W., Peng, M.-S., Zhang, Y.-P. (2022). African Suid Genomes Provide Insights into the Local Adaptation to Diverse African Environments. Molecular Biology and Evolution, 39(12). https://doi.org/10.1093/molbev/msac256.
Adeola, A. C., Bello, S., Abdussamad, A., Mark, A., Sanke, O., Onoja, A., Nneji, L., Abdullahi, N., Olaogun, S., Rogo, L., Mangbon, G., Pedro, S., Hiinan, M., Mukhtar, M., Ibrahim, J., Saidu, H., Dawuda, P., Bala, R., Abdullahi, H., Salako, A., Kdidi, S., Yahyaoui, M. H., Yin, T.-T. (2022). Scrapie-associated polymorphisms of the prion protein gene (PRNP) in Nigerian native goats. Gene, 855, 147121.
Liu, Y.-N., Chen, R.-M., Pu, Q.-T., Nneji, L., Sun, Y.-B. (2022). Expression Plasticity of Transposable Elements Is Highly Associated with Organismal Re-adaptation to Ancestral Environments. Genome Biology and Evolution, 14(6). https://doi.org/10.1093/gbe/evac084.
Adeola, A. C., Sola-Ojo, F., Opeyemi, Y., Oguntunji, A., Nneji, L., Ewuola, M., Bello, S., Olaniyi, W., Adesoji, A., Karuno, A., Sanke, O., Daniel, E. L. (2022). Genetic diversity and population structure of muscovy duck (Cairina moschata) from Nigeria. PeerJ. https://doi.org/10.7717/peerj.13236.
Oladipo, S., Nneji, L., Adeniyi, A., Adeyemi-Ale, O., Adelakun, K. (2022). Environmental correlates of some selected fish species in the Jebba Dam, North-Central Nigeria. Journal of Tropical Ecology, 38(3), 107–117. https://doi.org/10.1017/S0266467421000511.
Ugoh, A., Nneji, L., Adedeji, B., Adeyi, A. (2021). A preliminary survey of amphibians from the Idanre Forest Ecosystem, southwestern Nigeria. Journal of Wildlife and Biodiversity, 5(4), 1-7. https://doi.org/10.22120/jwb.2021.139825.1201.
Rahman, M. M., Nneji, L., Hossain, M. M., Nishikawa, K., Habib, K. A. (2021). Diversity and distribution of amphibians in central and northwest Bangladesh with an updated checklist for the country. Journal of Asia-Pacific Biodiversity, 15, 147-156. https://doi.org/10.1016/j.japb.2021.12.002.
Ayoola, A., Nneji, L., Odewumi, O., Oluwatobi, O., Emmanuel, S., Ogunjemite, B. (2021). Preliminary survey of non-human Primates in four forest areas of Yewa River Basin at the Nigeria-Benin Republic transboundary. Biologia, 77, 181–186. https://doi.org/10.1007/s11756-021-00891-x.
Jaynes, K., Myers, E., Gvoždík, V., Blackburn, D., Portik, D., Greenbaum, E., Jongsma, G., Rödel, M.-O., Badjedjea, G., Bamba-Kaya, A., Baptista, N., Akuboy, J., Ernst, R., Kouete, M., Kusamba, C., Masudi, F., McLaughlin, P., Nneji, L.,Onadeko, A., Penner, J., Vaz Pinto, P., Stuart, B., Tobi, E., Zassi-Boulou, A.-G., Leaché, A., Fujita, M., Bell, R. (2021). Giant Tree Frog diversification in West and Central Africa: Isolation by physical barriers, climate, and reproductive traits. Molecular Ecology, 31, 3979–3998. https://doi.org/10.1111/mec.16169.
Yan, F., Nneji, L., Jin, J.-Q. B., Yuan, Z.-y., Chen, J., Mi, X., Chen, H., Murphy, R., Che, J. (2021). Multi-locus genetic analyses of Quasipaa from throughout its distribution. Molecular Phylogenetics and Evolution, 163(107218). https://doi.org/10.1016/j.ympev.2021.107218.
Oladipo, S., Nneji, L., Iyiola, O., Nneji, I., Ayoola, A., Anifowoshe, A., Adelakun, K., Adeola, A., Mustapha, M. (2021). Patterns of ichthyofaunal diversity and distribution across Jebba Hydro-Electric Power (HEP) dam, Jebba, north-central Nigeria. Brazilian Journal of Biology. https://doi.org/10.1590/1519-6984.222952.
Nneji, L., Adeola, A., Adedeji, B., Olatunde, O., Onadeko, A., Eniang, E. A., Ayoola, A., Okeyoyin, A., Usongo, J., Adeyi, A., Oladipo, S., Rahman, M. M., Andem, A. B., Nneji, I., Ugwumba, O., Ugwumba, A., Adedeyi, A., Olory, C. S. (2021). Species richness, distribution pattern, and conservation status of amphibians in Cross River National Park, south‐eastern Nigeria. Biologia, 76, 2573–2588. https://doi.org/10.1007/s11756-021-00751-8.
Shen, Q.-K., Peng, M.-S., Adeola, A., Kui, L., Duan, S., Miao, Y.-W., Eltayeb, N., Lichoti, J., Otecko, N., Strillacci, M. G., Gorla, E., Bagnato, A., Charles, O., Sanke, O., Dawuda, P., Okeyoyin, A., Musina, J., Njoroge, P., Agwanda, B., Kusza, S., Nanaei, H. A., Peder, R., Xu, M.-M., Du, Y., Nneji, L., Murphy, R., Wang, M.-S., Esmailizadeh, A., Dong, Y., Ommeh, S., Zhang, Y.-P. (2021). Genomic Analyses Unveil Helmeted Guinea Fowl (Numida meleagris) Domestication in West Africa. Genome Biology and Evolution, 13(6). https://doi.org/10.1093/gbe/evab090.
Booth, H., Clark, M., Milner-Gulland, E.J., Amponsah-Mensah, K., Antunes, A. P., Brittain, S., Castilho, L., Campos-Silva, J., Constantino, P., Li, Y., Mandoloma, L., Nneji, L., Iponga, D., Moyo, B., McNamara, J., Rakotonarivo, S., Shi, J., Tagne, C., Velden, J., Williams, D. (2021). Investigating the risks of removing wild meat from global food systems. Current Biology, 31(8). https://doi.org/10.1016/j.cub.2021.01.079.
Oyebanji, O., Salako, G., Nneji, L., Oladipo, S., Bolarinwa, K.A., Chukwuma, E., Ayoola, A., Olagunju, T., Ighadalo, D., Nneji, I. (2020). Impact of climate change on the spatial distribution of endemic legume species of the Guineo-Congolian forest, Africa. Ecological Indicators, 122, 107282. https://doi.org/10.1016/j.ecolind.2020.107282.
Nneji, L., Adeola, A., Ayoola, A., Adedeji, B., Olatunde, O., Onadeko, A., Eniang, E. A., Ayoola, A., Oladipo, S., Wang, Y.-y., Mallan, Y., Anyaele, O., Nneji, I., Rahman, M. M., Olory, C. S. (2020). DNA barcoding and species delimitation of butterflies (Lepidoptera) from Nigeria. Molecular Biology Reports, 47, 9441–9457. https://doi.org/10.1007/s11033-020-05984-5.
Ayoola, A., Zhang, B.-L., Meisel, R., Nneji, L., Shao, Y., Morenikeji, O., Ayoola, A., Ng’ang’a, S., Ogunjemite, B., Okeyoyin, A., Ross, C., Wu, D.-D. (2020). Population Genomics Reveals Incipient Speciation, Introgression, and Adaptation in the African Mona Monkey (Cercopithecus mona). Molecular Biology and Evolution, 38(3), 876–890. https://doi.org/10.1093/molbev/msaa248.
Oladipo, S., Nneji, L., Anifowoshe, A., Nneji, I. (2020). Molecular characterization of Malapterurus minjiriya Sagua, 1987 and phylogenetic relationships within the genus Malapterurus (Silurifomes, Malapteruridae) from Nigerian inland water bodies. Journal of Fish Biology, 97, 1865–1869. https://doi.org/10.1111/jfb.14547.
Rahman, M. M., Nneji, L., Adeola, A., Chen, J., Eniang, E. A., Oladipo, S., Olatunde, O., Onadeko, A., Kilunde, F., Ayoola, A., Adedeji, B., Nneji, I., Akaowo, N., Ugwumba, A., Jin, J.-Q. B., Yin, T., Peng, M.-S., Olory, C. S., Eninekit, N., Che, J. (2020). Amphibian assemblages and diversity patterns in two forest ecosystems of South-Eastern Nigeria. African Journal of Ecology, 58, 815– 827. https://doi.org/10.1111/aje.12776.
Nneji, L., Adeola, A., Mustapha, M., Oladipo, S., Djagoun, C., Nneji, I., Adedeji, B., Olatunde, O., Ayoola, A., Okeyoyin, A., Ikhimiukor, O., Useni, G., Iyiola, O., Faturoti, E., Matouke, M., Ndifor, W., Wang, Y.-Y., Chen, J., Wang, W.-Z., Kachi, J., Ugwumba, O., Ugwumba, A., Nwani, C. (2020). DNA barcoding Silver butter catfish (Schilbe intermedius) reveals patterns of mitochondrial genetic diversity across African river systems. Scientific Reports, 10(7097). https://doi.org/10.1038/s41598-020-63837-4.
Adeola, A., Ewuola, M., Nneji, L., Oguntunji, A., Bello, S., Sola-Ojo, F., Ayoola, A., Adesoji, A., Sanke, O., Ebiakpor, D., Jonah, K., Jia, X., Wu, R.-N., Peng, M.-S., Zhang, Y.-P. (2020). Mitochondrial DNA variation of Nigeria Muscovy duck (Cairina moschata). Animal Genetics, 51, 485-486. https://doi.org/10.1111/age.12924.
Ajao, A., Nneji, L., Adeola, A., Oladipo, S., Ayoola, A., Wang, Y.-y., Adeniyi, A., Olademeji, Y. (2020). Genetic diversity and population structure of the native Western African honeybee (Apis mellifera adansonii Latreille, 1804) in Nigeria based on mitochondrial COI sequences. Zoologischer Anzeiger (Journal of Comparative Zoology), 293, 17-25. https://doi.org/10.1016/j.jcz.2021.05.007.
Nneji, L., Adeola, A., Wang, Y.-y., Ajao, A., Anyaele, O., Mallan, Y., Olatunde, O., Nneji, I., Ayoola, A., Rahman, M. M., Adeniyi, A., Okeyoyin, A., Olory, C. S. (2020). Testing the Effectiveness of DNA Barcoding for Biodiversity Assessment of Moths from Nigeria. Diversity, 12(2(85)). https://doi.org/10.3390/d12020085.
Nneji, L., Salako, G., Oladipo, S., Onadeko, A., Adedeji, B., Olatunde, O., Ugwumba, A., Adeola, A. (2019). Species Distribution Modelling predicts habitat suitability and reduction of suitable habitat under future climatic scenario for Sclerophrys perreti: A critically endangered Nigerian endemic toad. African Journal of Herpetology, 58, 481–491. https://doi.org/10.1111/aje.12713.
Ayoola, A., Ogunjemite, B., Nneji, L., Nkume, O. (2019). Comparative reproductive behavior in captive baboon (Papio anubis) and red capped mangabey (Cercocebus torquatus). Journal of Entomology and Zoology Studies, 7(6), 479-483. http://www.entomoljournal.com.
Nneji, L., Adeola, A., Okeyoyin, A., Oladipo, O., Saidu, Y., Samuel, D., Usongo, J., Adedeji, B., Olatunde, O., Adeyi, A., Ugwumba, O., Ugwumba, A. (2019). Diversity and distribution of amphibians and reptiles in Gashaka Gumti National Park, Nigeria. Herpetology Notes, 12, 543-559.
Yang, C., Fu, T., Lan, X., Zhang, Y., Nneji, L., Murphy, R., Sun, Y., Che, J. Comparative Skin Histology of Frogs Reveals High-elevation Adaptation of the Tibetan Nanorana Parkeri. Asian Herpetological Research, 10(2), 79-85. https://doi.org/10.16373/j.cnki.ahr.180093.
Nneji, L., Adeola, A., Yan, F., Okeyoyin, A., Oladipo, O., Saidu, Y., Samuel, D., Nneji, I., Olagunju, T., Olatunde, O., Oladipo, S., Iyiola, O., Auta, T., Usman, A., Abdullahi, H., Ikhimiukor, O., Zhou, W.-W., Jin, J.-Q., Adiaha, O., Ugwumba, A., Peng, M.-S., Murphy, R., Che, J. (2019). Genetic variation and cryptic lineage diversity of the Nigerian red-headed rock agama Agama agama associate with eco-geographic zones. Current Zoology, 65(6), 713–724. https://doi.org/10.1093/cz/zoz002.
Nneji, L., Adeola, A., Okeyoyin, A., Onadeko, A., Eniang, E. A., Adeyi, A., Oladipo, S., Adedeji, B., Usongo, J., Andem, A. B., Ugwumba, O., Ugwumba, A., Olory, C. S. (2019). First record of Foulassi screeching frog Arthroleptis adelphus Perret 1966 (Anura, Arthroleptidae) from Nigeria, with notes on its phylogenetic position. Check List, 15(1), 253–258. https://doi.org/10.15560/15.2.253.
Nneji, L., Adeola, A., Okeyoyin, A., Oladipo, O., Saidu, Y., Usongo, J., Ugwumba, A. (2019). Assessing the effectiveness of molecular data for species identification and diversity studies of Nigerian herpetofauna. Mitochondrial DNA Part B Resources, 4(1), 713–724. https://doi.org/10.1080/23802359.2018.1561219.
Ayoola, A., Wang, Y., Nneji, L., Adeola, A., Ogunjemite, B., Dong-Dong, W. (2019). Complete mitochondrial genome sequence for the Cercopithecus erythrotis camerunensis (Primate: Cercopithecidae). Mitochondrial DNA Part B, 4(1), 209-210. https://doi.org/10.1080/23802359.2018.1546138.
Oladipo, S., Nneji, L., Anifowoshe, A., Nneji, I., Iyiola, O., Adeola, A., Mustapha, M. (2018). Growth Pattern and Condition Factor of Seven Commercially Important Freshwater Fish Species of Jebba Lake, North-Central Nigeria. Iranian Journal of Ichthyology, 5(2), 167-172. https://doi.org/10.22034/iji.v5i2.272.
Iyiola, O., Nneji, L., Mustapha, M., Nzeh, C., Oladipo, S., Nneji, I., Okeyoyin, A., Nwani, C., Ugwumba, O., Ugwumba, A., Faturoti, E., Wang, Y.-Y., Chen, J., Wang, W.-Z., Adeola, A. (2018). DNA barcoding of economically important fish species of North – Central Nigeria uncovers cryptic lineages. Ecology and Evolution, 8(14), 6932–6951. https://doi.org/10.1002/ece3.4210.
Adeyi, A. O., Nneji, L., Adeyi, E., Somade, O., Agbaogun, B. (2018). Combined use of earthworm (Alma millsoni) and bacterium (Bacillus sp.) improved the bioremediation of spent engine oil contaminated soil. Chemistry and Ecology, 34(6), 532–547. https://doi.org/10.1080/02757540.2018.1472247.
Nneji, L., Adeola, A., Yan, F., Okeyoyin, A., Oladipo, O., Olagunju, T., Olatunde, O., Oladipo, S., Iyiola, O., Auta, T., Usman, A., Abdullahi, H., Peng, M.-S., Murphy, R., Jin, J.-Q., Ugwumba, A., Che, J. (2017). Cryptic diversity of Nigerian Agama within West African Radiation. Russian Journal of Herpetology, 25(2), 97-112. http://rjh.folium.ru/index.php/rjh/article/view/1303.
Adeola, A., Oluwole, O., Oladele, B., Olorungbounmi, T., Boladuro, B., Olaogun, S., Nneji, L., Sanke, O., Dawuda, P., Omitogun, O., Frantz, L., Murphy, R., Xie, H.-B., Peng, M.-S., Zhang, Y.-P. (2017). Analysis of the genetic variation in mitochondrial DNA, Y-chromosome sequences, and MC1R sheds light on the ancestry of Nigerian indigenous pigs. Genetics, Selection, and Evolution, 49(52). https://doi.org/10.1186/s12711-017-0326-1.
Nneji, L., Somade, O., Adeyi, A. O. (2016). Earthworm-assisted bioremediation of petroleum hydrocarbon-contaminated soils from motorcar mechanic workshops in Ibadan, Oyo State, Southwestern Nigeria. Bioremediation Journal, 20(4), 263–285. https://doi.org/10.1080/10889868.2016.1250723.